 - 1. Maxilla (Mx) with a maxillary brush (MxB) and the cardo (Cd) fused with the base of the palpus
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 - 2. Hypostomal suture (HyS) complete
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 - 3. Occipital foramen (OF) circular with a distinct collar
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 - 4. Seta 8-M absent
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 - 5. Pecten (Pt) absent
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 - 6. Siphon (S) without midventral filaments
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