This information reflects views on the classification of scorpions, based
on Dr. Stockwell's Ph.D. dissertation, “Revision of the phylogeny and
higher classification of scorpions (Chelicerata)”, from 1989. The supra-generic
classification is sequenced and annotated. However, the order of the listing
of the genera for each category does not imply any phylogenetic relationship.
Fossil taxa are marked by a dagger (†). The cladograms on
which this classification are based are also available.
Class SCORPIONIDA Latreille, 1810
SCORPIONIDA incertae sedis:
†Genus Brontoscorpio Kjellesvig-Waering, 1972
†Genus Feistmantelia Fric, 1904
†Genus Bromsgroviscorpio Kjellesvig-Waering, 1986
†Genus Titanoscorpio Kjellesvig-Waering, 1986
†Genus Wattisonia Wills, 1960
†Genus Palaeomachus Pocock, 1911
†Genus Tiphoscorpio Kjellesvig-Waering, 1986
†Genus Eskiscorpio Kjellesvig-Waering, 1986
†Genus Microlabis Corda, 1839
†Genus Hydroscorpius Kjellesvig-Waering, 1986
†Order PROTOSCORPIONES Petrunkevich, 1949:
- †Genus Palaeophonus Thorell and Lindström, 1884
- †Genus Allopalaeophonus Kjellesvig-Waering, 1986
- †Genus Dolicophonus Petrunkevitch, 1949
- †Genus Palaeoscorpius Lehmann, 1944
†Order PALAEOSCORPIONES, new:
-
†Superfamily Proscorpioidea Scudder, 1885:
- †Genus Proscorpius Whitfield, 1885
- †Genus Archaeophonus Kjellesvig-Waering, 1966
- †Genus Waeringoscorpio Størmer, 1970
- †Genus Labriscorpio Leary, 1980
- †Genus Stoermeroscorpio Kjellesvig-Waering, 1986
-
†Superfamily Archaeoctonoidea Petrunkevitch,
1949:
- †Genus Archaeoctonus Pocock, 1911
- †Genus Pseudoarchaeoctonus Kjellesvig-Waering, 1986
- †Genus Loboarchaeoctonus Kjellesvig-Waering, 1986
Order SCORPIONES Hemprich & Ehrenberg, 1837:
SCORPIONES incertae sedis:
- †Genus Praearcturus Woodward, 1871
- †Genus Branchioscorpio Kjellesvig-Waering, 1986
- †Genus Acanthoscorpio Kjellesvig-Waering, 1986
- †Genus Gigantoscorpio Størmer, 1963
- †Genus Centromachus Thorell and Lindström, 1885
- †Genus Cyclophthalmus Corda, 1835
- †Genus Anthracochaerilus Kjellesvig-Waering, 1986
- †Genus Trachyscorpio Kjellesvig-Waering, 1986
- †Genus Scoloscorpio Kjellesvig-Waering, 1986
- †Genus Garnettius Petrunkevitch, 1953
- †Genus Palaeobuthus Petrunkevitch, 1913
- †Genus Boreoscorpio Kjellesvig-Waering, 1986
- †Genus Leioscorpio Kjellesvig-Waering, 1986
- †Genus Opsieobuthus Kjellesvig-Waering, 1986
- †Genus Pseudobuthiscorpius Kjellesvig-Waering, 1986
- †Genus Petaloscorpio Kjellesvig-Waering, 1986
- †Genus Waterstonia Kjellesvig-Waering, 1986
- †Genus Stenoscorpio Kjellesvig-Waering, 1986
- †Genus Willsiscorpio Kjellesvig-Waering, 1986
- †Genus Spongiophonus Wills, 1947
- †Genus Liassiscorpionides Bode, 1951
- †Genus Mioscorpio Kjellesvig-Waering, 1986
†Suborder MESOSCORPIONINA, new:
- †Genus Mesophonus Wills, 1910
- †Genus Mazonia Meek and Worthen, 1868
- †Genus Heloscorpio Kjellesvig-Waering, 1986
- †Genus Phoxiscorpio Kjellesvig-Waering, 1986
- †Genus Isobuthus Fric, 1904
- †Genus Eobuthus Fric, 1904
- †Genus Eoscorpius Meek and Worthen, 1868
- †Genus Pareobuthus Wills, 1959
- †Genus Kronoscorpio Kjellesvig-Waering, 1986
- †Genus Paraisobuthus Kjellesvig-Waering, 1986
- †Genus Telmatoscorpio Kjellesvig-Waering, 1986
Suborder NEOSCORPIONINA Thorell & Lindström,
1885:
-
†Infraorder PALAEOSTERNI, new:
- †Genus Eoctonus Petrunkevitch, 1913
- †Genus Buthiscorpius Petrunkevitch, 1953
- †Genus Allobuthiscorpius Kjellesvig-Waering, 1986
- †Genus Apiscorpio Kjellesvig-Waering, 1986
- †Genus Anthracoscorpio Kusta, 1885
- †Genus Lichnoscorpius Petrunkevitch, 1949
- †Genus Allobuthus Kjellesvig-Waering, 1986
- †Genus Coseleyscorpio Kjellesvig-Waering, 1986
-
Infraorder ORTHOSTERNI Pocock, 1911:
-
ORTHOSTERNI incertae sedis:
- †Genus Palaeopisthacanthus Petrunkevitch, 1913
- †Genus Compsoscorpius Petrunkevitch, 1949
-
Superfamily Buthoidea Simon, 1879:
-
Family Chaerilidae Pocock,
1893:
- Genus Chaerilus Simon, 1877
-
Family Buthidae Simon,
1879:
- Genus Akentrobuthus Lamoral, 1976
- Genus Ananteris Thorell, 1891
- Genus Microbuthus Kraepelin, 1898
- Genus Butheolus Simon, 1883
- Genus Butheoloides Hirst, 1925
- Genus Orthochirus Karsch, 1892
- Genus Buthus Leach, 1815
- Genus Androctonus Hemprich and Ehrenberg, 1829
- Genus Leiurus Hemprich and Ehrenberg, 1829
- Genus Hemibuthus Pocock, 1900
- Genus Mesobuthus Vachon, 1950
- Genus Darchenia Vachon, 1977
- Genus Anomalobuthus Kraepelin, 1901
- Genus Apistobuthus Finnegan, 1932
- Genus Babycurus Karsch, 1886
- Genus Birulatus Vachon, 1974
- Genus Buthacus Birula, 1908
- Genus Buthiscus Biruls, 1905
- Genus Odontobuthus Vachon, 1950
- Genus Odonturus Karsch, 1879
- Genus Kraepelinia Vachon, 1974
- Genus Liobuthus Birula, 1898
- Genus Lissothus Vachon, 1948
- Genus Cicielus Vachon, 1948
- Genus Compsobuthus Vachon, 1949
- Genus Psammobuthus Birula, 1911
- Genus Hoplocystis Karsch, 1884
- Genus Hottentotta Birula, 1908
- Genus Vachoniolus Levy, Amitai & Shulov, 1973
- Genus Vachonus Tikader & Bastewade, 1983
- Genus Isometrus Hemprich & Ehrenberg, 1829
- Genus Isometroides Keyserling, 1885
- Genus Lychas Koch, 1845
- Genus Lychasoides Vachon, 1974
- Genus Pocockius Francke, 1985
- Genus Pseudolychas Kraepelin, 1912
- Genus Scorpiobuthus Werner, 1939
- Genus Charmus Karsch, 1879
- Genus Karasbergia Hewitt, 1914
- Genus Parabuthus Pocock, 1890
- Genus Uroplectes Peters, 1862
- Genus Grosphus Simon, 1880
- Genus Tityobuthus Pocock, 1893
- Genus Centruroides Marx in Howard, 1890
- Genus Rhopalurus Thorell, 1876
- Genus Tityus Koch, 1836
- Genus Mesotityus González-Sponga, 1982
- Genus Tityopsis Armas, 1974
- Genus Alayotityus Armas, 1973
- Genus Microtityus Kjellesvig-Waering, 1966
- Genus Zabius Thorell, 1894
-
Superfamily Chactoidea Pocock, 1893:
-
Family Chactidae Pocock,
1893:
- Genus Nullibrotheas Williams, 1974
- Genus Chactas Gervais in Walckenaer & Gervais, 1844
- Genus Teuthraustes Simon, 1878
- Genus Vachoniochactas González-Sponga, 1978
- Genus Chactopsis Kraepelin, 1912
- Genus Brotheas Koch, 1837
- Genus Broteochactas Pocock, 1893
-
Family Euscorpiidae Laurie,
1896:
- Genus Euscorpius Thorell, 1876
- Genus Megacormus Karsch, 1881
- Genus Plesiochactas Pocock, 1900
- Genus Troglocormus Francke, 1982
-
Family Scorpiopsidae Kraepelin,
1899:
- Genus Scorpiops Peters, 1862
- Genus Parascorpiops Banks, 1928
- Genus Neoscorpiops Vachon, 1980
- Genus Euscorpiops Vachon, 1980
- Genus Alloscorpiops Vachon, 1980
- Genus Dasyscorpiops Vachon, 1974
-
Superfamily Scorpionoidea Peters, 1862:
-
Family Bothriuridae Simon,
1880:
-
Subfamily Lisposominae Lawrence, 1928:
- Genus Lisposoma Lawrence, 1928
-
Subfamily Bothriurinae Simon, 1880:
- Genus Thestylus Simon, 1880
- Genus Phoniocercus Pocock, 1893
- Genus Tehuankea Cekalovic, 1973
- Genus Brachistosternus Pocock, 1893
- Genus Orobothriurus Maury, 1975
- Genus Bothriurus Peters, 1862
- Genus Timogenes Simon, 1880
- Genus Vachonia Abalos, 1954
- Genus Urophonius Pocock, 1893
- Genus Centromachetes Lönnberg, 1897
- Genus Cercophonius Peters, 1862
-
Family Ischnuridae Simon,
1879:
-
Subfamily Heteroscorpioninae Birula, 1903:
- Genus Heteroscorpion Birula, 1903
-
Subfamily Hemiscorpiinae Pocock, 1893:
- Genus Hemiscorpius Peters, 1862
- Genus Habibiella Vachon, 1974
-
Subfamily Ischnurinae Simon, 1879:
- Genus Hadogenes Kraepelin, 1894
- Genus Chiromachetes Pocock, 1899
- Genus Cheloctonus Pocock, 1892
- Genus Opisthacanthus Peters, 1862
- Genus Chiromachus Pocock, 1893
- Genus Iomachus Pocock, 1893
- Genus Liocheles Sundevall, 1833
-
Family Urodacidae Pocock,
1893:
- Genus Urodacus Peters, 1862
-
Family Scorpionidae Peters,
1862:
- Genus Scorpio Linnaeus, 1758
- Genus Opistophthalmus Koch, 1837
- Genus Pandinus Throell, 1876
- Genus Heterometrus Hemprich and Ehrenberg, 1829
-
Family Diplocentridae Pocock,
1893:
-
Subfamily Diplocentrinae Pocock, 1893:
- Genus Diplocentrus Peters, 1962
- Genus Didymocentrus Kraepelin, 1905
- Genus Bioculus Stahnke, 1968
- Genus Tarsoporosus Francke, 1978
- Genus Cazierius, Francke, 1978
- Genus Oiclus Simon, 1880
- Genus Heteronebo Pocock, 1899
-
Subfamily Nebinae Kreapelin, 1905:
- Genus Nebo Kraepelin, 1898
-
Superfamily Vaejovoidea Thorell, 1876:
-
Family Vaejovidae Thorell,
1876:
-
Subfamily Uroctoninae Mello-Leitão, 1934:
- Genus Uroctonus Thorell, 1976
- Genus Uroctonites Williams & Savary 1991
-
Subfamily Vaejovinae Thorell, 1876:
- Genus Vaejovis Koch, 1836 (Mexicanus & Granulatus species
groups)
-
Subfamily Syntropinae Kraepelin, 1905:
-
New Tribe:
- Nitidulus species group
-
Tribe Syntropini Kraepelin, 1905:
- Genus Serradigitus Stahnke, 1974
- Genus Syntropis Kraepelin, 1901
- Punctipalpi Species Group
- Eusthenura Species Group
-
Tribe Pseudouroctonini, provisional:
- Genus Pseudouroctonus Stahnke, 1974
-
Tribe Paruroctonini, provisional:
- Genus Paravaejovis Williams, 1980
- Genus Paruroctonus Werner, 1934
- Genus Smeringurus Haradon, 1983
- Genus Vejovoidus. Stahnke, 1974
-
Family Superstitionidae Stahnke,
1940:
-
Subfamily Superstitioninae Stahnke, 1940:
- Genus Superstitionia Stahnke, 1940
-
Subfamily Troglotayosinae, new:
- Genus Troglotayosicus Lourenço, 1981
-
Subfamily Belisariinae, new:
- Genus Belisarius Simon, 1879
-
Subfamily Typhlochactinae Mitchell, 1968:
- Genus Alacran Francke, 1982
- Genus Sotanochactas Francke, 1986
- Genus Typhlochactas Mitchell, 1968
-
Family Iuridae Thorell,
1876:
-
Subfamily Iurinae Thorell, 1876:
- Genus Iurus Thorell, 1876
- Genus Paraiurus Francke, 1986
-
Subfamily Caraboctoninae Kraepelin, 1905:
- Genus Caraboctonus Pocock, 1893
- Genus Hadruroides Pocock, 1893
-
Subfamily Hadrurinae Stahnke, 1974:
- Genus Hadrurus Thorell, 1876
- Genus Anuroctonus Pocock, 1893